One of the most confusing aspects of the Nikon flash system is this thing called 'Auto FP High Speed Sync'. This article will hopefully clear up the mystery about this mode.
Before you can understand FP High Speed Sync, you first need to understand what FP means and how the Normal Flash Sync works.
Normal Flash Sync
FP stands for Focal Plane and it refers to the type of shutter used in most modern DSLR cameras. A focal plane shutter is actually two precisely timed curtains positioned between the lens and the sensor that can either block light from hitting the sensor or allow light to hit the sensor. The reason there are two shutter curtains is to be able to get much higher effective shutter speeds.
It is important to understand that these curtains open and close in exactly the same amount of time. So the shutter speed is set by timing between the start of the first curtain opening and the start of the second curtain closing.
Notice that the entire sensor will be open to the light at every shutter speed up to the speed of the curtain movement itself. This is the maximum normal Flash Sync Speed. To say this in another way; at all speeds up to the maximum normal Flash Sync Speed, the first curtain completely opens before the second curtain begins to close. At any shutter speed higher than this, the second curtain will begin closing before the first curtain gets fully open, thus never exposing the entire sensor at any one time. At really high shutter speeds, this results in very narrow 'slit' of light that travels across the sensor.
These two curtains travel vertically across the opening from the top to the bottom, and this in itself causes some strange effects when using high shutter speeds (small slit). If you shoot something that moves horizontally really fast like a race car from the side, you can sometimes see that the wheels seem to lean forward a bit because the top part of the wheel was exposed after the bottom part and the top moved forward a little bit as the slit moved from bottom to top. Of course, the image is inverted on the focal plane, exposing from the bottom to the top of the image, and that's why race car wheels lean forward even though the shutter moves from top to bottom. Also, this effect only occurs if you don't pan with the car. You can totally eliminate the forward leaning effect by panning.
As mentioned, a focal plane shutter mechanism moves the curtains at a very precise speed. This speed is determined during manufacturing of the mechanism and is governed mostly by how recently the shutter was designed. In older 35mm cameras, this speed was 1/60th second, but with time, shutters got faster and faster, and in the new D300 this speed is 1/320th sec. And those race car wheels lean much further forward with an older 1/60th shutter than with the D300 1/320th shutter.
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